Collective Hysteresis and Multistability in Threshold Networks

Abstract

In a mechanistic model of the dawn chorus, Kaye showed that heterogeneous activation thresholds and a shared feedback signal determined by the population's active fraction can produce abrupt collective activation and hysteresis. We extend this mechanism to a network of interacting agents. Each node has a continuous activation level, and a nonnegative row-stochastic matrix determines how node activities contribute to individual feedback. We prove that sufficiently weak feedback yields a unique globally attracting equilibrium. For all feedback strengths, the homogeneous dynamics exactly reproduce Kaye's scalar equation; consequently, network topology does not alter the folds or cusp of the homogeneous branch, and no heterogeneous mode becomes unstable before the homogeneous mode. For equitable partitions, the network admits an exact quotient system in which nodes within a block receive the same aggregate input from every block. When blocks are uncoupled, the quotient reduces to independent copies of Kaye's scalar equation. We show that every assignment of stable scalar equilibria to blocks persists under sufficiently weak interblock coupling, producing a combinatorial family of stable quotient equilibria that lift to stable full-network equilibria and remain under small perturbations that break exact equitability. For two symmetrically coupled blocks, branches with unequal block activities terminate at a pair of symmetry-related cusp bifurcations. Near the onset of bistability, we derive scaling laws for the interblock coupling at which these bifurcations occur, the activity difference between the blocks at bifurcation, and the corresponding shift of the external stimulus from the scalar cusp. Numerical continuation confirms the scaling laws for gamma, logistic, and normal threshold distributions.